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“Multiagency collaboration is widely used in contemporary service delivery systems. This article explores the interpersonal interactions within collaborative systems, among subsystems, and among organizations. [The authors’] focus is on illuminating the informal mechanisms that facilitate collaboration, joint production, coordination and integration of service delivery, and sustained effort. … [The authors] use a multiple case-study approach to explore the dynamics of informal accountability among individuals working within county-based children’s service systems in three states [and] find informal interpersonal dynamics nested in combinations of vertical and horizontal ties with mixed administrative authority arrangements derived from both formal and informal accountability relationships. These data reveal shared norms, facilitative behaviors, informal rewards and sanctions, and challenges that create the dynamics of informal accountability. Informal accountability is shaped by the prevalence of relationship building and champion behavior as facilitative behaviors, discernible tension between the operation of formal and informal accountability systems, a gap between the rhetoric of collaboration and the reality of collaborative service provision, differences in informal accountability dynamics across hierarchical levels within service delivery systems, and the critical roles of street-level caseworkers in informal accountability.”